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Climate models have evolved into Earth system models with representation of the physics, chemistry, and biology of terrestrial ecosystems. This companion book to Gordon Bonan's Ecological Climatology: Concepts and Applications, Third Edition, builds on the concepts introduced there, and provides the mathematical foundation upon which to develop and understand ecosystem models and their relevance for these Earth system models. The book bridges the disciplinary gap among land surface models developed by atmospheric scientists; biogeochemical models, dynamic global vegetation models, and ecosystem demography models developed by ecologists; and ecohydrology models developed by hydrologists. Review questions, supplemental code, and modeling projects are provided, to aid with understanding how the equations are used. The book is an invaluable guide to climate change and terrestrial ecosystem modeling for graduate students and researchers in climate change, climatology, ecology, hydrology, biogeochemistry, meteorology, environmental science, mathematical modeling, and environmental biophysics.